& date of birth :
Rhodos, Greece, 18 May 1941
Via G Pascoli 60, 20133 Milan, Italy
Position : Full professor of Biology and Genetics, University Vita-Salute, Via Olgettina 60, 20135 Milan, Italy
Graduated in Physics, University of Florence.
and scientific career
CNR research position, International Institute of Genetics and Biophysics,
CNR senior scientist, Center for Molecular and Cellular Pharmacology,
Associate Professor of Population Genetics, Faculty of Science,
University of Naples.
Associate Professor of Human Genetics, First Medical School, University
1996-2001: Professor of Genetics & Biology, University Vita-Salute, Milan.
2000-: Full Professor Genetics and Biology.
2001-2004: Director, SISSA-ISA, International School for Advanced Studies, Trieste
2005: EMBO Award for Communication in the Life Sciences
EMBO (1988), Academia Europaea (1995), HUGO (1997), EMBO Council
Editorial Board: Mechanisms of Development (1990-), EMBO Journal (1993-1995), Experimental Cell Research (1994-2000), Human Molecular Reproduction (1995-2000), Gene (1996-2005), NeuroPsychoanalysis (2000-), BMC Neuroscience (2000-), Lingue e Linguaggio (2002-)
Postgraduate work on two-quanta radiation processes.
Genetic analysis of rDNA at the bobbed locus of D. melanogaster.
Genetics and Biochemistry of position effect in D. melanogaster.
Molecular investigation of the internal repeats of the fruit fly rDNA
Organization of human rDNA. Non canonical Class I HLA genes in normal and
The human HOX homeobox gene family. Structural and temporal colinearity.
Sequential activation of HOX genes.
Isolation and characterization of Evx, Dlx, Otx and Emx homeobox genes.
Role of Otx genes in early mouse, frog, chick and zebrafish embryos and in
1997-2001: Emx genes in the developing cerebral cortex. Emx2 mutations in the human
congenital cortical defect schizencephaly. Emx2 in proliferation, migration and
regionalisation of the cerebral cortex.
2001-: Cognitive Neuroscience and Public Understanding of Science.